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Cross-talks and perspectives Both GSH and Pro, with molecular formula C 10 H 17 N 3 O 6 S and C 5 H 9 NO 2 , respectively, belong to the glutamate or -ketoglutarate family and originate from a common precursor L-glutamate (Moat et al., GSH and Pro may occur in the concentrations of few mM (23 mM) in various plant tissues (Noctor et al., via N and S nutrition, respectively (Anjum et al., 2 s 1 ) (Ogawa et al., 2 O 2 (or exogenous H 2 O 2 ) can increase Pro level by modulating the ex-novo synthesis of Pro (Matysik et al., via GSH reductase (Anjum et al., Both GSH (Ogawa, via metabolizing the varied ROS and their reaction products (Ruiz and Blumwald, A differential coordination of other components of ascorbate (AsA)-GSH pathway (enzymes such as ascorbate peroxidase, GSH reductase, GSH peroxidase, GSH sulfo-transferase, monodehydroascorbate reductase, dehydroascorbate reductase and catalase