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J Biol Chem 275:3145131459 Vatamaniuk OK, Mari S, Lang A et al (2004) Phytochelatin synthase, a dipeptidyltransferase that undergoes multisite acylation with -glutamylcysteine during catalysis: stoichiometric and site-directed mutagenic analysis of Arabidopsis thaliana PCS1-catalyzed phytochelatin synthesis